This fourth installment of "The Year in Evolutionary Biology" series includes reviews on competition and the evolutionary dynamics of mutualism; rapid Hox gene evolution and the evolution of body plans; the evolutionary and conservation consequences of inbreeding-environment interactions; evolution and development of character displacement; applications of information theory to evolution; and gene duplications.

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Timothy A. Mousseau is the editor of The Year in Evolutionary Biology 2012, Volume 1251, published by Wiley. Charles W. Fox is the editor of The Year in Evolutionary Biology 2012, Volume 1251, published by Wiley.